Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands
BACKGROUND AND OBJECTIVES: Odonate larvae play an important role in macroinvertebrate trophic networks and are excellent indicators of wetland quality. However, despite their ecological importance, research on odonates and how they interact with their environment is scarce. This study aims to assess...
- Autores:
-
Moreno Pallares, María Inés
Bonilla, Maria Argenis
Guillot Monroy, G.H.
Torregroza Espinosa, Ana Carolina
- Tipo de recurso:
- Article of journal
- Fecha de publicación:
- 2023
- Institución:
- Corporación Universidad de la Costa
- Repositorio:
- REDICUC - Repositorio CUC
- Idioma:
- eng
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- oai:repositorio.cuc.edu.co:11323/10323
- Acceso en línea:
- https://hdl.handle.net/11323/10323
https://repositorio.cuc.edu.co/
- Palabra clave:
- Odonata larvae
Diversity
Limnology
Predator
Tropical wetlands
- Rights
- openAccess
- License
- Atribución 4.0 Internacional (CC BY 4.0)
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dc.title.eng.fl_str_mv |
Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands |
title |
Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands |
spellingShingle |
Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands Odonata larvae Diversity Limnology Predator Tropical wetlands |
title_short |
Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands |
title_full |
Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands |
title_fullStr |
Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands |
title_full_unstemmed |
Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands |
title_sort |
Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands |
dc.creator.fl_str_mv |
Moreno Pallares, María Inés Bonilla, Maria Argenis Guillot Monroy, G.H. Torregroza Espinosa, Ana Carolina |
dc.contributor.author.none.fl_str_mv |
Moreno Pallares, María Inés Bonilla, Maria Argenis Guillot Monroy, G.H. Torregroza Espinosa, Ana Carolina |
dc.subject.proposal.eng.fl_str_mv |
Odonata larvae Diversity Limnology Predator Tropical wetlands |
topic |
Odonata larvae Diversity Limnology Predator Tropical wetlands |
description |
BACKGROUND AND OBJECTIVES: Odonate larvae play an important role in macroinvertebrate trophic networks and are excellent indicators of wetland quality. However, despite their ecological importance, research on odonates and how they interact with their environment is scarce. This study aims to assess macroinvertebrate composition as a determinant of larval abundance in Miathyria marcella (Odonata: Anizoptera: Libellulidae). METHODS: In total, 29 samples were collected from six wetlands with different types of hydrological influence using standardized invertebrate sampling techniques in Atlántico, a department located in northern Colombia. Standardized invertebrate sampling techniques were used at 29 sampling points. Obtained data were used to analyze invertebrate abundance and a non-parametric multidimensional scaling analysis was applied. In addition, a correlation analysis was conducted between macroinvertebrate composition and Miathyria marcella larval abundance. FINDINGS: A total of 2586 larvae and 12925 individual macroinvertebrates were collected, distributed across 25 orders and 58 families. The most abundant orders were Neotaenioglossa (26 percent), Odonata (15 percent) Calanoida (10 percent) and Diptera (8 percent). Heatmap and scaling analysis indicated different macroinvertebrate compositions in the sampled wetlands. It was found a high positive correlation between Miathyria marcella and the orders Odonata (R2 = 0.84, p-value ≤ 0.05), Coleoptera (R2 = 0.52, p-value ≤ 0.05), Basommatophora (R2 = 0.60, p-value ≤ 0.05), and Hemiptera (R2 = 0.50, p-value ≤ 0.05). CONCLUSION: The results suggest that the abundance of Miathyria marcella responds to the accompanying macroinvertebrates, the composition of which depends on the type of hydrological influence. Assessment approaches that focus on the relationships between macroinvertebrate taxa are important conservation tools for biodiversity assessment. Results from this study will serve as a baseline to propose monitoring and follow-up strategies for environmental sustainability in wetlands in this region. |
publishDate |
2023 |
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2023-07-21T21:01:04Z |
dc.date.available.none.fl_str_mv |
2023-07-21T21:01:04Z |
dc.date.issued.none.fl_str_mv |
2023 |
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Artículo de revista |
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http://purl.org/coar/resource_type/c_2df8fbb1 |
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dc.identifier.citation.spa.fl_str_mv |
Moreno Pallares, M., Bonilla Gómez, M., Guillot Monroy, G., Torregroza-Espinosa, A. Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands. Global Journal of Environmental Science and Management, 2023; 9(1): 129-140. doi: 10.22034/gjesm.2023.01.10 |
dc.identifier.issn.spa.fl_str_mv |
2383-3572 |
dc.identifier.uri.none.fl_str_mv |
https://hdl.handle.net/11323/10323 |
dc.identifier.doi.none.fl_str_mv |
10.22034/gjesm.2023.01.10 |
dc.identifier.eissn.spa.fl_str_mv |
2383-3866 |
dc.identifier.instname.spa.fl_str_mv |
Corporación Universidad de la Costa |
dc.identifier.reponame.spa.fl_str_mv |
REDICUC - Repositorio CUC |
dc.identifier.repourl.spa.fl_str_mv |
https://repositorio.cuc.edu.co/ |
identifier_str_mv |
Moreno Pallares, M., Bonilla Gómez, M., Guillot Monroy, G., Torregroza-Espinosa, A. Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands. Global Journal of Environmental Science and Management, 2023; 9(1): 129-140. doi: 10.22034/gjesm.2023.01.10 2383-3572 10.22034/gjesm.2023.01.10 2383-3866 Corporación Universidad de la Costa REDICUC - Repositorio CUC |
url |
https://hdl.handle.net/11323/10323 https://repositorio.cuc.edu.co/ |
dc.language.iso.spa.fl_str_mv |
eng |
language |
eng |
dc.relation.ispartofjournal.spa.fl_str_mv |
Global Journal of Environmental Science and Management |
dc.relation.references.spa.fl_str_mv |
Abdul, N.H.; Rawi, C.S.M.; Ahmad, A.H.; Al-Shami, S.A., (2017). Effect of environmental disturbances on odonata assemblages along a tropical polluted river. Ekológia., 36(4): 388-402 (15 pages). Akram, W.; Ali-Khan, H.A., (2016). Odonate nymphs, generalist predators and their potential in the management of dengue mosquito, Aedes aegypti (Diptera: Culicidae). J. ArthropodBorne Dis., 10(2): 252-257 (6 pages). Barbán, L., (2015). Odonata larvae associated with the root system of Eichhornia crassipes (pontederiaceae), in the chalons dam, Santiago de Cuba, Cuba. Revista Colombiana De Ciencia Animal Recia., 7(2): 130-138 (9 pages). Bowles, D.E.; Kleinsasser, L.J., (2022). Environmental determinates of distribution for dragonfly nymphs (Odonata: Anisoptera) in Urban and Non-Urban East Texas Streams, USA. Hydrobiology., 1(1): 76-88 (13 pages). Buczynska, E.; Buczynski, P., (2019). Survival under anthropogenic impact: the response of dragonflies (Odonata), beetles (Coleoptera) and caddisflies (Trichoptera) to environmental disturbances in a two-way industrial canal system (central Poland). PeerJ., 6: e6215 (31 pages). Bybee, S.M.; Kalkman, V.J.; Erickson, R.J.; Fradsen, P.B.; Breinholt, J.W.; Suvorov, A.; Dijkstra, K.B.; Cordero-Rivera, A.; Skevington, J.H.; Abbott, J.C.; Sanchez, M.; Lemmon, A.R.; Moriarty, E.; Ware, J.L., (2021). Phylogeny and classification of Odonata using targeted genomics. Mol. Phylogenet. Evol., 160: 107115 (15 pages). Carvalho-Soares, A.A.; Ferreira, K.G.; Sousa, K.S.; Nascimento, A.C.L.; Mendoza- Penagos, C.C.; Vieira, T.B.; Salcedo, A.K.M.; Oliveira-Junior, J.M.B.; Calvão, L.B.; Dias-Silva, K., (2022). Checklist and New Occurrences of Odonata (Insecta) from Volta Grande do Xingu, Pará, Brazil. Hydrobiology., 1(2): 183-195 (13 pages). Castillo, R.M.; Huamantinco, A.A., (2020). Variación espacial de la comunidad de macroinvertebrados acuáticos en la zona litoral del humedal costero Santa Rosa, Lima, Perú. Rev. Biol. Trop., 68(1): 50-68 (19 pages). Choi, J.; Kim, S.; Kim, J.; Kwon, S., (2020). Habitat Preferences and Trophic Position of Brachydiplax chalybea flavovittata Ris, 1911 (Insecta: Odonata) Larvae in Youngsan River Wetlands of South Korea. Insects., 11(5): 1-18 (18 pages). Choo, M.Z.J.; Low, B.W.; Ngiam, R.W.J.: Yeo, D.C.J., (2021). Predation of mosquitos by odonates in a tropical urban environment: insights from functional response and field mesocosm experiments. Biol. Control. 161(104702): 1-9 (9 pages). Cid, N.; Bonada, N.; Carlson, S.M.; Grantham, T.E.; Gasith, A.; Resh, V.H., (2017). High variability is a defining component of Mediterranean-climate rivers and their biota. Water., 9(52): 1-24 (24 pages). Correa-Araneda, F.; Núñez, D.; Díaz, M.E.; Gómez-Capponi, F.; Figueroa, R.; Acuña, J.; Boyero, L.; Esse, C., (2021). Comparison of sampling methods for benthic macroinvertebrates in forested wetlands. Ecol. Indic., 125(107551): 1-11 (11 pages). Czerniawska-Kusza, I., (2022). Assessing benthic macroinvertebrates in relations to environmental variables and revitalisation works. Ecol. Chem. Eng. S., 29(1): 99-110 (12 pages). Dou, Q.; Du, X.; Cong, Y.; Wang, L.; Zhao, C.; Song, D.; Liu, H.; Huo, T., (2022). Influence of environmental variables on macroinvertebrate community structure in Lianhuan Lake. Ecol. Evol., 12(2): 1-15 (15 pages). Fulan, J.Â.; Anjos, M., (2015). Predation by Erythemis nymphs (Odonata) on Chironomidae (Diptera) and Elmidae (Coleoptera) in different conditions of habitat complexity. Acta Limnol. Bras., 27(4): 454-458 (5 pages). Gómez-Tolosa, M.L.; Mendoza-Cuenca, L.F.; Rioja-Paradela, T.M.; Espinoza-Medinilla, E.E.; Alonso-Eguía, P.E.; Rivera-Velázquez, G.; Penagos-García, F.E.; Pérez-Munguía, R.M.; Ortega-Salas, H.; Gómez-Cristiani, M.; Gómez-Gutiérrez, R.B., (2015). Odonata (Insecta) de tres cuencas en la costa de Chiapas: lista de especies y registro nuevo. Rev. Mex. Biodivers., 86(1): 1-7 (7 pages). González-Soriano, E.; Noguera, F.A.; Pérez-Hernández, C.X.; ZaragozaCaballero, S.; González-Valencia, L., (2021). Patterns of richness, diversity and abundance of an odonate assemblage from a tropical dry forest in the Santiago Dominguillo Region, Oaxaca, México (Insecta: Odonata). Biodivers. Data J., 9(e60980): 1-21 (21 pages). Guellaf, A.; Bennas, N.; El Haissoufi, M.; L’Mohdi, O.; Kettani, K., (2021). New data on the biodiversity and chorology of aquatic insects (Odonata, Coleoptera and Hemiptera) of Martil Basin (northwestern Morocco). Graellsia. 77(2): 1-23 (23 pages). Hamada, N.; Thorp, J.H.; Rogers, D.C., (2018). Keys to Neotropical Hexapoda. Thorp and Covich’s Freshwater Invertebrates. Vol. 3, Academis Press, Elsevier. Hecca, D.; Arinafril, A.; Novia, N., (2018). Diversity of odonata and aquatic environmental conditions in lake area (water ski and opi) Jakabaring Palembang-South Sumatera. Biovalentia. 4(2): 1-6 (6 pages). Karbassi, A.R.; Pazoki, M., (2015). Environmental qualitative assessment of rivers sediments. Global J. Environ. Sci. Manage., 1(2): 109-116 (8 pages). Kondo, T.; Palacino-Rodriguez, F.; Pena-Cuellar, R.D., (2015). Report of Erpetogomphus sabaleticus Williamson, 1918 feeding on Diaphorina citri Kuwayama (Hemiptera: Liviidae). Boletin del Museo de entomologia., 16(1): 17-26 (10 pages). Mandal, S.K.; Ghosh, A.; Bhattacharjee, I.; Chandra, G., (2018). Biocontrol efficiency of odonate nymphs against larvae of the mosquito, Culex quinquefasciatus Say, 1823. Acta Tropica., 106(2): 109-114 (6 pages). Mariani-Ríos, A.; Maldonado-Benítez, N.; Ramírez, A., (2022). Natural history of Odonata assemblages in tropical streams in Puerto Rico. Neotro. Biodivers., 8(1): 112-123 (12 pages). May, M.L., (2019). Odonata: Who They Are and What They Have Done for Us Lately: Classification and Ecosystem Services of Dragonflies. Insects., 10(62): 1-17 (17 pages). Ramírez, A.; Gutiérrez-Fonseca, P.E., (2014). Functional feeding groups of aquatic insect families in Latin America: a critical analysis and review of existing literature. Rev. Biol. Trop., 62(2): 155-167 (13 pages). Rivera-Usme, J.; Pinilla, G.; Rangel-Churio, J.; Castro, M.; CamachoPinzón, D., (2015). Biomass of macroinvertebrates and physicochemical characteristics of water in an Andean urban wetland of Colombia. Braz. J. Biol., 75(1): 180-190 (11 pages). Rychla, A.; Benndorf, J.; Buczynski, P., (2011). Impact of pH and conductivity on species richness and community structure of dragonflies (Odonata) in small mining lakes. Fundam. Appl. Limnol., 179(1): 41-50 (10 pages). Samanmali, C.; Udayanga, L.; Ranathunge, T.; Perea, S.; Hapugoda, M.; Weliwitiya C., (2018). Larvicidal potential of five selected dragonfly nymphs in Sri Lanka over Aedes aegypti (Linnaeus) larvae under laboratory settings. BioMed. Res. Int., 8759459: 1-10 (10 pages). Schwentner, M.; Timms, B.V.; Richter, S., (2015). Spinicaudata (Branchiopoda: Diplostraca) in australia’s arid zone: unparalleled diversity at regional scales and within water bodies. J. Crust Biol., 35(3): 366-378 (13 pages). Silva, E.; Rocha, A.; Leal, M.; Santos, O.; Sousa, J.; Viana, A.R.; Silva, K.K.; Matos, E.; Santos, E.; Landim, A.; Gimenez, T., (2020). Freshwater mollusks from three reservoirs of Piauí, northeastern Brazil. Biota Neotropica. 20(1): 1-8 (8 pages). Suárez-Morales, E., (2015). Chapter 29 - Class Maxillopoda, in: Thorp, J. H., Rogers, D. C. (Eds.), Thorp and Covich’s Freshwater Invertebrates (4th. Edition). Academic Press (47 pages). Tonkin, J.D.; Bogan, M.T.; Bonada, N.; Rios-Touma, B.; Lytle, D., (2017). Seasonality and predictability shape temporal species diversity. Ecology., 98(5): 1201-1216 (16 pages). Torres-Bejarano, F.M.; Torregroza-Espinosa, A.C.; Martinez-Mera, E.; Castaneda-Valbuena, D.; Tejera-Gonzalez, M.P., (2020). Hydrodynamics and water quality assessment of a coastal lagoon using environmental fluid dynamics code explorer modeling system. Global J. Environ. Sci. Manage., 6(3): 289-308 (20 pages). Vilenica, M., (2017). Ecological traits of dragonfly (Odonata) assemblages along an oligotrophic Dinaric karst hydrosystem. Ann Limnol., 53: 377-389 (13 pages). Younes, A.; El-Sherif, H.; Gawish, F.; Mahmoud, M., (2015). Potential of Hemianax ephippiger (Odonata-Aeshnidae) nymph as predator of Fasciola intermediate host, Lymnaea natalensis. Asian Pac. Trop. Biomed., 5(8): 671-675 (5 pages). |
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© Journal Management System. Powered by ejournalplus. |
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Atribución 4.0 Internacional (CC BY 4.0) |
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Atribución 4.0 Internacional (CC BY 4.0) © Journal Management System. Powered by ejournalplus. https://creativecommons.org/licenses/by/4.0/ http://purl.org/coar/access_right/c_abf2 |
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Atribución 4.0 Internacional (CC BY 4.0)© Journal Management System. Powered by ejournalplus.https://creativecommons.org/licenses/by/4.0/info:eu-repo/semantics/openAccesshttp://purl.org/coar/access_right/c_abf2Moreno Pallares, María InésBonilla, Maria ArgenisGuillot Monroy, G.H.Torregroza Espinosa, Ana Carolina2023-07-21T21:01:04Z2023-07-21T21:01:04Z2023Moreno Pallares, M., Bonilla Gómez, M., Guillot Monroy, G., Torregroza-Espinosa, A. Macroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands. Global Journal of Environmental Science and Management, 2023; 9(1): 129-140. doi: 10.22034/gjesm.2023.01.102383-3572https://hdl.handle.net/11323/1032310.22034/gjesm.2023.01.102383-3866Corporación Universidad de la CostaREDICUC - Repositorio CUChttps://repositorio.cuc.edu.co/BACKGROUND AND OBJECTIVES: Odonate larvae play an important role in macroinvertebrate trophic networks and are excellent indicators of wetland quality. However, despite their ecological importance, research on odonates and how they interact with their environment is scarce. This study aims to assess macroinvertebrate composition as a determinant of larval abundance in Miathyria marcella (Odonata: Anizoptera: Libellulidae). METHODS: In total, 29 samples were collected from six wetlands with different types of hydrological influence using standardized invertebrate sampling techniques in Atlántico, a department located in northern Colombia. Standardized invertebrate sampling techniques were used at 29 sampling points. Obtained data were used to analyze invertebrate abundance and a non-parametric multidimensional scaling analysis was applied. In addition, a correlation analysis was conducted between macroinvertebrate composition and Miathyria marcella larval abundance. FINDINGS: A total of 2586 larvae and 12925 individual macroinvertebrates were collected, distributed across 25 orders and 58 families. The most abundant orders were Neotaenioglossa (26 percent), Odonata (15 percent) Calanoida (10 percent) and Diptera (8 percent). Heatmap and scaling analysis indicated different macroinvertebrate compositions in the sampled wetlands. It was found a high positive correlation between Miathyria marcella and the orders Odonata (R2 = 0.84, p-value ≤ 0.05), Coleoptera (R2 = 0.52, p-value ≤ 0.05), Basommatophora (R2 = 0.60, p-value ≤ 0.05), and Hemiptera (R2 = 0.50, p-value ≤ 0.05). CONCLUSION: The results suggest that the abundance of Miathyria marcella responds to the accompanying macroinvertebrates, the composition of which depends on the type of hydrological influence. Assessment approaches that focus on the relationships between macroinvertebrate taxa are important conservation tools for biodiversity assessment. Results from this study will serve as a baseline to propose monitoring and follow-up strategies for environmental sustainability in wetlands in this region.12 páginasapplication/pdfengGJESM PublicationIranhttps://www.gjesm.net/article_253268.htmlMacroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlandsArtículo de revistahttp://purl.org/coar/resource_type/c_6501http://purl.org/coar/resource_type/c_2df8fbb1Textinfo:eu-repo/semantics/articlehttp://purl.org/redcol/resource_type/ARTinfo:eu-repo/semantics/publishedVersionhttp://purl.org/coar/version/c_970fb48d4fbd8a85Global Journal of Environmental Science and ManagementAbdul, N.H.; Rawi, C.S.M.; Ahmad, A.H.; Al-Shami, S.A., (2017). Effect of environmental disturbances on odonata assemblages along a tropical polluted river. Ekológia., 36(4): 388-402 (15 pages).Akram, W.; Ali-Khan, H.A., (2016). Odonate nymphs, generalist predators and their potential in the management of dengue mosquito, Aedes aegypti (Diptera: Culicidae). J. ArthropodBorne Dis., 10(2): 252-257 (6 pages).Barbán, L., (2015). Odonata larvae associated with the root system of Eichhornia crassipes (pontederiaceae), in the chalons dam, Santiago de Cuba, Cuba. Revista Colombiana De Ciencia Animal Recia., 7(2): 130-138 (9 pages).Bowles, D.E.; Kleinsasser, L.J., (2022). Environmental determinates of distribution for dragonfly nymphs (Odonata: Anisoptera) in Urban and Non-Urban East Texas Streams, USA. Hydrobiology., 1(1): 76-88 (13 pages).Buczynska, E.; Buczynski, P., (2019). Survival under anthropogenic impact: the response of dragonflies (Odonata), beetles (Coleoptera) and caddisflies (Trichoptera) to environmental disturbances in a two-way industrial canal system (central Poland). PeerJ., 6: e6215 (31 pages).Bybee, S.M.; Kalkman, V.J.; Erickson, R.J.; Fradsen, P.B.; Breinholt, J.W.; Suvorov, A.; Dijkstra, K.B.; Cordero-Rivera, A.; Skevington, J.H.; Abbott, J.C.; Sanchez, M.; Lemmon, A.R.; Moriarty, E.; Ware, J.L., (2021). Phylogeny and classification of Odonata using targeted genomics. Mol. Phylogenet. Evol., 160: 107115 (15 pages).Carvalho-Soares, A.A.; Ferreira, K.G.; Sousa, K.S.; Nascimento, A.C.L.; Mendoza- Penagos, C.C.; Vieira, T.B.; Salcedo, A.K.M.; Oliveira-Junior, J.M.B.; Calvão, L.B.; Dias-Silva, K., (2022). Checklist and New Occurrences of Odonata (Insecta) from Volta Grande do Xingu, Pará, Brazil. Hydrobiology., 1(2): 183-195 (13 pages).Castillo, R.M.; Huamantinco, A.A., (2020). Variación espacial de la comunidad de macroinvertebrados acuáticos en la zona litoral del humedal costero Santa Rosa, Lima, Perú. Rev. Biol. Trop., 68(1): 50-68 (19 pages).Choi, J.; Kim, S.; Kim, J.; Kwon, S., (2020). 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Biomed., 5(8): 671-675 (5 pages).14012919Odonata larvaeDiversityLimnologyPredatorTropical wetlandsPublicationORIGINALMacroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands.pdfMacroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands.pdfArtículoapplication/pdf830444https://repositorio.cuc.edu.co/bitstreams/4a77ca1a-c741-4c9a-b90c-b98b5f78e688/download7a73c40230bcb15254ca7e7459bee0a3MD51LICENSElicense.txtlicense.txttext/plain; charset=utf-814828https://repositorio.cuc.edu.co/bitstreams/5af2d1dc-e880-47d5-8a01-10ab40a48883/download2f9959eaf5b71fae44bbf9ec84150c7aMD52TEXTMacroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands.pdf.txtMacroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands.pdf.txtExtracted texttext/plain41139https://repositorio.cuc.edu.co/bitstreams/debb6260-91c0-42bc-89af-467959467f74/downloadace5d111270e09ad575837fffadc2b28MD53THUMBNAILMacroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands.pdf.jpgMacroinvertebrates composition as determinants of larval abundance in the dragonfly Miathyria marcella in tropical wetlands.pdf.jpgGenerated Thumbnailimage/jpeg12094https://repositorio.cuc.edu.co/bitstreams/823dd2cb-0c79-4c90-bf06-81b4698b99dc/downloadb4ebccc9306214f99f95dbd5c0f3ba77MD5411323/10323oai:repositorio.cuc.edu.co:11323/103232024-09-17 12:47:44.218https://creativecommons.org/licenses/by/4.0/© Journal Management System. 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CUCrepdigital@cuc.edu.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